Map of Santander Bay (Cantabrian Sea, North-Atlantic coast of Spain) showing the three sampling sites: HI Horadada Island, MP Magdalena Peninsula, and MI Mouro Island
In the littoral rocky bottoms of the Santander Bay (Fig. 1 ),
hydrodynamic and topographic variables produce a patchy
distribution of benthic macrohabitats (García-Castrillo et al.
The study was carried out at three different sites: Mouro Island
(MI, number of quadrants ( n ) = 294), Magdalena
Peninsula (MP, n = 29), and Horadada Island
(HI, n = 34), located outside, at the entrance,
and inside of the Bay of Santander (Atlantic coast of northern
Spain; Fig. 1 ), respectively.
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